Nocturnal and Flier are also important, although with Mystcraft ages or regular sleeping you won't absolutely need them. Cave-dwelling can be useful but it's far from essential.
Edit: Some quick tests show that bees consider the times 12550-23450 to be night. Meaning a standard bee will only work about 54.6% of a full day/night cycle, so Nocturnal can raise productivity by up to 83%.
Flier is much less predictable but will most likely be between 10-30%, rainmakers and Thaumcraft flux can however make this vary wildly.
Again all of these numbers will be reduced significantly with frequent sleeping, or completely negated by eternal day/no weather Mystcraft ages.