At the moment who the hell knows, the rate that bee addon mods are updating/changing & little info that follows is a hindrance, you learn a branch next update some names have changed/steps added or removed/breeding items requirements changed, it would really help if there was some more documentation regarding the magic & extra changes.
That said in the past I have found around 5 species that will only breed in a required biome (apart from the normal ones we all know of like nether) or produce there specialties, so most all bees are fine in normal/normal with +2 both, if after half a stack of drone you dont get the specialty then its time to start looking for reasons.